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Tough call, maybe throughout their lifespan Capcom are number 1 but my vote would have to go to Rare. This may be a silly argument but whilst Capcom and Square were huge for Nintendo they were also big in helping other rival companies (mainly Sony) which knocked Nintendo into 2nd place during the 5th gen.

Rare produced great (and mostly exclusive) games just for Nintendo and were vitally important in stopping the N64 from being a total flop.