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The biggest change is the lighting. That is night and day better than Smash 4. Animations are are vastly improved. Textures, shaders, and overall level of detail are also higher, but not as improved as the lighting and animations. And most models probably have a somewhat higher polygon count, but that is probably the least significant upgrade.

Of course Dark Samus would see a huge improvement. Your model is going to get a lot more attention when it goes from assist trophy to full-fledged playable character.