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curl-6 said:

Without spoiling anything, I do like that the game makes the effort to give even the villains some backstory, so there's more to them than just "I like being evil."

Honestly, I think characterisation in this game is the best in the series.  Characters have dimension and emotional range and growth.  Every villain has a motive and something to generate empathy.  Or at least pitty.  The game even made me like a Nopon, which I though was impossible.