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Nuvendil said:

Or those were preliminary character designs that were later scrapped?  It's also entirely possible that's a test build that uses test assets, especially character models since they are rather inconsequentual in the context of game mechanics testing.  Main character designs in a game like this can come pretty late in development and can also change drastically over time. 

I for one, *highly* doubt that it has main-character co-op because it would be a huge hurdle for the narrative based on what we have seen so far of it.  And this is Takahashi we're talking about; he's not going to hamstring his narrative.

Highly doubtful. This gameplay footage was shown over 3 years after development started, and just a few months before E3 2014 at which it was fully unveiled, main characters and all. If that's not enough, there's this:

Elma, right? This picture is from the January 2013 reveal trailer.

I don't see why main character co-op would represent a 'hurdle' for the narrative. Cross is a member of the military, other players' main characters are simply other members of the military.