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Hard to remember them all right now but I think these are my top 5 moments so far this gen:

 

5.   No More Heroes  (Wii):  The fifth ranked assassin boss "fight" (Henry's first appearance).

4.   Metal Gear Solid 4  (PS3):  The "Beauty and the Beast" corps' first appearance ("Show me your rage!")

3.   Uncharted 2  (PS3):  The whole helicopter chase and collapsing building sequence (actually that whole level is great).

2.   Metal Gear Solid 4  (PS3):  The end of act 3; starting with the motorcycle chase and continuing to the end cinematic where liquid takes control (wonderfully directed).

1.   Bioshock  (X360/PS3):  Three words:  "Would you kindly?" (just amazing) and the reveal of Atlus' true identity.  I mean seriously, few movies or books have plot twists of this magnitude.

 

Cool thread btw.