To be honest for the majority of the game I got by with just upgrading radomly. General rules are to upgrade using animal bits to give you bonuses and then upgrade with machine parts. I know that if you want the "ultimate" weapons that any starting weapon can be upgraded to becoma an ultimate weapon but will have different stats depending on what you start with. For instance, if you start with a Gladius and upgrade/transform to Omega weapon your strength will be V.high (+750) whilst your magic bonus will only be +200 or something. If you upgrade starting with Lionheart it'll be about +500 for each.
Although you're pretty far into the story the upgrading only really takes off towards the very end. I think by Chpt. 12 I'd maxed out the first level (before first transformation) of most of the characters weapons.
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