@Khuutra
Basically what nordlead said: Gnome and KDE come each with their slightly different approach in interfaces and attitude. The KDE camp is generally more about very powerful and configurable applications. The Gnome camp is generally more about offering less customization -or hiding it behind admin-level tools- and simpler experiences.
A lot of fanboy silliness ensues, sadly, but they both offer very valid environments.
You can totally use KDE applications such as Amarok in Ubuntu's Gnome, or install KDE and run Gnome apps such as rythmbox in it. The only downside (besides slight memory and HDD penalties, as I said) is that you might notice inconsistencies e.g. in button ordering (Gnome orders button Cancel-Ok a-la-Mac ; KDE orders them Ok-Cancel a-la-Windows) or in how other UI elements behave.
If you don't care about those littleendians vs bigendians disputes (you have bigger inconsistencies in windows between the media player, office and other applications) then you can totally forget the Gnome/KDE thing :) If you're an interface consistency freak, on the other hand, you might want to stick with applications of one camp only.
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