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The thing about MGS4 is that it is much more segmented than any of the previous MGS games. It's more or less like playing separate episodes, complete with "intermissions" during inter-act installations (my one and only real gripe with MGS4). I would have preferred to see the 4GB install limit ditched and just have all 5 acts ready, regardless of HDD space.

Each act varies so much that it is almost like playing in a different game every time with the same game engine/inventory. The game engine, inventory system and camo system have all been refined from MGS3 although the basic mechanics are pretty similar (MGS4 still plays the smoothest in terms of controls, etc. by a wide margin IMO)

And yes, everything from MGS3 ties in with MGS4, particularly Act 3 and the conclusion.