Soriku said:
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In simple terms, the protagonist is the main character, while the antagonist is the character who opposes the main character. Not all stories have a true villain, but I suggest a simple definition of a villain in a work of fiction is a character whose actions are condemned by the narrative.
It is very much possible for the protagonist to be a villain, and the antagonist to be the "hero." See also the police in GTA -- there is nothing villainous about them whatsoever, whereas the protagonist is sometimes the true villain (but sometimes just a sort of antihero).