I'm not sure on the whole 1 million polygons per car thing, as I have only heard it from other people, but I would guess that a lot of heavy optimisation goes into displaying the game. In most games, the further away an object gets from the camera, the less detail needs to be displayed. Developers will deform the models and reduce the poly count the further away an object is, whilst making sure there is no noticable drop in quality. Models can therefore be changed to have half, a quarter or even less the polygons than the original based on distance. For some closeups like viewing the Model while tuning, or getting that view of the car wheel at the start of a race, it probably has the full 1 million (if that is correct) as its viewing only a section of the full model... but certainly when racing, the figure for each car would be well below that figure. The type of optimisations would happen in GT5 as well...
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