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LBP may be the one.

PS3 is doing pretty damn well for not having a defining title (outselling 360 over the same amount of time).  It will be interesting to see what happens if it gets one.

On the other hand, I don't think Sony consoles generally have had them as much as others (Mario, Sonic, Halo).  Sony has a huge number of compelling 1st party IPs, but none alone seem to define a system.